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在非洲游历了一圈之后，纽约中央公园的四位明星们继续着回到纽约，回到中央公园的旅程。不过，只要旅途中有猩猩和那群企鹅的“帮忙”，旅程就没有想象中的那么顺利。在蒙特卡洛的一家赌场里，由于大家的耍宝，大闹赌场，进而招致了心狠手辣的动物管理局上尉杜波依斯的围追堵截。为了逃过追捕，大家逃进了一个在欧洲进行巡回演出的马戏团。在马戏团里，大家延续着“马达加斯加”式的幽默，笑料百出。虽然，欧洲这一路上的旅行并不顺利，不断受到杜波依斯上尉的追捕威胁，不过大家一直怀揣着回到纽约，回到纽约中央公园的梦想。在马戏团，还帮助了东北虎维塔利、美洲虎吉亚以及海狮斯蒂芬诺重拾对马戏表演的信心。。伊森·亨特带领IMF小队继续与AI智体对抗，打响终极之战。。Chorus girls Polly, Carol and Trixie are ecstatic when they learn that Broadway producer Barney Hopkins is putting on a new show. He promises all of the girls parts in the new show and even hires their neighbor Brad Roberts, an unknown composer, to write some of the music. There's only one problem: he doesn't have the money to bankroll it all. That problem is solved when Brad turns out to be quite rich but he insists that he not perform. When opening night comes, the juvenile lead can't go on forcing Brad to take the stage. He's recognized of course and his upper crust family wants him to quit. When he refuses, they tell him to end his relationship with Polly or face having his income cut off. When Brad's snobbish brother Lawrence mistakes Carol for Polly, the girls decide to have a bit of fun and teach him a lesson.。